It's hard to tell for certain because it is difficult for me to reproduce the 
bug.
I think that it would usually be between nautilus and thunar.

I can always reproduce this, slightly different, bug:
* Right click on a file in Nautilus and click copy.
* Open a new copy of Thunar
* Right click and notice that the "Paste" command is grayed out.

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Can't copy + paste files between windows (quite often)
https://bugs.launchpad.net/bugs/231923
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