I have a very similar problem. Account 1 uses SMTP 1, and account 2 SMTP
2. However whenever I want to reply from account 2 it uses SMTP 1, and
unsuccessfully. I have to remove the message from the queue. Changing
the default account in prefs changes the behavior so that SMTP 2 is
always used rather than SMTP 1.

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evolution uses wrong smtp server when replying to mail
https://bugs.launchpad.net/bugs/21989
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