It comes up for me as well from time to time after a few weeks of a
fresh install. Though it seems to start the theme after a couple of
seconds, however nautilus remains at the broken theme.

The problem seems to be that 'gnome-settings-daemon' crashes, and nautilus 
starts before it's up again.
It can be fixed with 'nautilus -q' once 'gnome-settings-daemon' is running 
stable. I'll post the crash report as soon as i catch it. But apart from 
gnome-settings-daemon being crashed, it should force nautilus also to reload.
Cheers

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https://bugs.launchpad.net/bugs/521766

Title:
  Theme not applied on startup

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