This is a very ambiguous topic and I suspect it is actually two or more bugs.
However, it seems to me that one fix (mentioned bygynoid, kingaaronj, ttp and
which also helped me) is definitive and would be as follows.
When choosing a default sound device in gnome-sound-properties, it
should also change the default PulseAudio device as does the
padevchooser package.
** Also affects: gnome-control-center
Importance: Undecided
Status: New
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9.04 flash player no sound after upgrade
https://bugs.launchpad.net/bugs/367173
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