I see the same faulty behaviour in gnome-volume-control, meaning the "Toggle 
audio recording from Capture" always is off (the state with the white X on red 
ground on it) when opening the window.
I can't reproduce it right now, but it seemed like the state of that setting 
had some effect as to whether I could record something on my system or not, one 
or two hours ago today. But after also playing around a little with alsamixer, 
the microphone now always works, regardless of this setting.

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Can't toggle audio recording for devices in Volume Control applet to 'on' state
https://bugs.launchpad.net/bugs/290543
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