I happen to read this thread 3 years ago and got interested in Prof. Jim's 
statement that " It was also asked about computing heritability “for each 
landmark”. That would be difficult without some strong assumptions."

I understand there has been discussion in this forum about the 
inappropriateness of analysis of landmarks one at a time after GPA. 
However, I am curious to learn under what "strong assumption" could we 
perform analysis "one landmark at a time"?

Shape by definition is the configuration of ALL landmarks after GPA. This 
means one should never analyze landmarks one at a time with the slightest 
hope that Procrustes shape coordinates for a single landmark represent 
local shape information. I think understanding the exceptional condition 
where one could analyze landmarks one at a time helps me gain deeper 
understanding of this issue.

Thank you.

Kieran

On Wednesday, July 31, 2019 at 9:49:38 PM UTC+8 [email protected] wrote:

> Dear Prof. Houle,
>
> Thank you very much for your interest in my questions. My dataset contains 
> 3D body shape data of people from peopole of two ethnicities. Based on such 
> phenotypic data, I wish to estimate population differentiation and 
> selection differential/gradient, and total predicted response to selection. 
> I agree with you that estimation of these quantities requires phenotypic 
> and additive genetic variance-covariance matrix rather than heritability. I 
> have only phenotypic data at the moment, although I will obtain the genetic 
> data sometime later.
>
> I understand that one approach to estimate the G matrix is to multiply the 
> P matrix by an estimated value of heritability. Implicit in this approach 
> is the assumption of proportionality. Am I right that it is impossible to 
> estimate G in the absence of genetic data, apart from the approach above?
>
> Regards,
> Patrick
>
> On Wednesday, 31 July 2019 03:10:31 UTC+8, dhoule wrote:
>
>> First of all, quantitative genetic concepts are defined relative to a 
>> reference or base population.  If you are mixing individuals from a wide 
>> geographic area, what is your population that you are making inferences 
>> about?
>>
>> Second, you can't do any quantitative genetics unless you have both 
>> 'unrelated' (relative to the defined reference population), and individuals 
>> with known degrees of relationship.   Do you have genomic data on these 
>> individuals? 
>>
>> If all you have are individuals of unknown relationship, all you can do 
>> is calculate the P matrix, then make some very strong assumption - like 
>> heritability is 0.3, and get a sort of proxy G matrix.  This might be 
>> better than nothing.
>>
>> Why do you want to know the heritability?  To predict the magnitude of 
>> changes,  you need the genetic variance and covariances, rather than 
>> heritabilities.  
>>
>> David Houle
>> On 7/30/2019 11:59 AM, lv xiao wrote:
>>
> A huge thank you to all who have contributed to my questions. It did took 
>> me a while to read the articles kindly suggested. I now have some basic 
>> understanding of heritability, selection differential, selection gradient, 
>> etc. 
>>
>> I have 3D data on body shape of Europeans and Africans and I wish to 
>> obtain estimates of heritability and visualize shape changes associated 
>> with selection differential and response to selection. The first thing I 
>> would like to do is to estimate the additive genetic variace-covariance 
>> matrix G, and the phenotypic variance-covariance matrix P. Software such as 
>> ACE and Wombat have been used for such purpose. The essence of this seems 
>> to be a multivariate mixed-effect model where the dependent variable is the 
>> shape data in matrix and the independent variables consist of both fixed 
>> and random effects. However, my dataset is not related to full-sibling 
>> design and I would like to ask how could I derive matrix P and G from my 
>> phenotypic data? 
>>
>> Once these two matrices are estimated, I will be able to performe further 
>> analyses on my own. Thank you.
>>
>> On Wednesday, 24 July 2019 21:12:13 UTC+8, lv xiao wrote: 
>>>
>>> Dear all, 
>>>
>>> Heritability is traditionally estimated based on genetic data. However, 
>>> it has also been estimated from phenotypic data. In this case, twin 
>>> comparison is the common study design. My question is if I have a sample of 
>>> *indenpendent 
>>> individuals* whose 3D shape data are available, is it possible to 
>>> *calculate 
>>> heritability based on Procrustes shape coordinates* for overall shape 
>>> and for each landmark?
>>>
>>> Several studies have used the Sequential Oligogenic Linkage Analysis 
>>> Routines (SOLAR) package for estimation of heritability of craniofacial 
>>> trait based on independent samples (J. Anat.(2009) 214, pp19–35; THE 
>>> ANATOMICAL RECORD 298:1535–1547 (2015)). However, these traits are linear 
>>> distances, which are quite different from "shape" based on Procrustes shape 
>>> coordinates. 
>>>
>>> Thanks
>>>
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>>
>> -- 
>> David Houle
>> Department of Biological Science
>> Florida State University
>> 319 Stadium Drive
>> Tallahassee, FL 32306-4295
>> 850-339-5670 <(850)%20339-5670>http://bio.fsu.edu/~dhoule/
>>
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