Using bisection, I've determined that this was introduced in the
following xenial commit:

commit 69747b70c731cc7081e7d13a0f536bb451f5a385
Author: Iyappan Subramanian <isubraman...@apm.com>
Date:   Mon Jul 25 17:12:42 2016 -0700

    drivers: net: xgene: Enable MDIO driver

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https://bugs.launchpad.net/bugs/1722579

Title:
  xgene nic unresponsive after ifdown/ifup

Status in linux package in Ubuntu:
  Fix Released
Status in linux source package in Xenial:
  In Progress

Bug description:
  [Impact]
  Sean Feole discovered that if you ifdown/ifup the NIC on an X-Gene 2 system, 
the NIC does not come back online. While the interface is reported as up, you 
cannot ping the system. In addition to the obvious problems this causes for a 
sysadmin, it also breaks software that does this automatically, such as juju 
when it sets up a host bridge.

  [Test Case]
  Boot an X-Gene 2 system, and start pinging it from a remote system. Assuming 
the pinged interface is eth0, run:

  $ sudo ifdown eth0; sudo ifup eth0

  System will cease responding to pings.

  [Regression Risk]
  (Still under investigation)

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