While investigating this failure, I noticed the following output in the test log:
> library('glmmTMB') Warning message: In checkMatrixPackageVersion() : Package version inconsistency detected. TMB was built with Matrix version 1.2.18 Current Matrix version is 1.3.2 Please re-install 'TMB' from source using install.packages('TMB', type = 'source') or ask CRAN for a binary version of 'TMB' matching CRAN's 'Matrix' package This is a clear sign that r-cran-tmb is missing some versioned dependency on the version of r-cran-matrix that it was built against. I did try rebuilding r-cran-tmb locally and indeed, the warning message did go away, but unfortunately this did not help with the actual failure.